In 1926 Italian mathematician and physicist, Vito Volterra of Sapienza – Università di Roma known for his contributions to mathematical biology, published "Varizioni e fluttuazioni del numero d'individui in specie animali conviventi" in Mem. R. Acad. Naz. dei Lincei (ser.6) II, 31-113. In this paper Volterra created the basic equations for two species interactions.
This work was translated into English and published in the journal Nature the same year as "Fluctuations in the abundance of a species considered mathematically".